Gram-Positive Bacteria | Your Medical Advices

Tuesday, September 9, 2014

Gram-Positive Bacteria


General Characteristics

        -Gram-positive bacteria have a highly cross-linked, multilayered (and usually thick) peptidoglycan    cell wall which traps the large Gram crystal violet-iodine complex staining them deep purple.

        -Teichoic acids are linked to either the cytoplasmic membrane (the lipoteichoic acids) or to the cell wall peptidoglycan. Teichoic acids are unique to Gram-positive bacteria and play roles in adherence and triggering Gram-positive shock as the cell wall is broken down.
        -A variety of cell surface proteins are present, which are organism specific.
        -Gram-positive bacteria have no outer membrane and therefore no hydrophobic barrier to limit access of larger antibiotics to the peptidoglycan.

            Major genera of Gram-positive bacteria


            • Genus: Staphylococcus. Common name: Staphylococci
              • Features
                • Gram-positive cocci generally in tight grapelike clusters or, in specimens, as singlets, pairs, or short chains as well as clusters.
                • Catalase-positive, breaking down hydrogen peroxide into water and oxygen.
                • Facultative anaerobes producing energy more efficiently aerobically.
                • Haloduric (salt-tolerant).
                • Speciated medically on the basis of coagulase and hemolysis.
            • Genus: Streptococcus. Common name: Streptococci
              • Features
                • Gram-positive cocci in chains or pairs. They tend to be oval.
                • Distinguished from staphylococci on the basis of the catalase test. Streptococci are catalase-negative, aerotolerant anaerobes that grow in full oxygen but ferment both in the presence and absence of oxygen.
                • Subdivided or speciated by three different systems: serology, hemolysin production, and biochemical properties.
                  • Serology using Lancefield’s antibodies to cell wall carbohydrates
                    • Streptococci that are positive for these carbohydrates are classified into Lancefield’s serogroups (e.g., Group A strep). There are now more than 20 groups.
                    • Bacteria that have these cell wall carbohydrates produce a pyogenic reaction.
                    • Some alpha-hemolytic streptococci including Strep. pneumoniae and the viridans streptococci lack these cell wall carbohydrates. They are not grouped using Lancefield’s antibodies and are not pyogenic.
                  • Hemolysin testing
                  • Biochemical tests
                • Streptococci are mainly opportunists but can cause disease in debilitated patients or if they gain entry into the body.
            • Genus: Enterococcus
              • Features
                • Catalase-negative, facultative anaerobes fermenting even in full oxygen; Streptococcal family.
                • Alpha-hemolytic or nonhemolytic, Gram-positive cocci in chains that have the Group D streptococcal cell wall carbohydrate.
                • Part of the normal human gastrointestinal (GI) flora.
                • Tolerant of high concentrations of bile salts and NaCl.
                • Have a high level of drug resistance that continues to increase due to efficient acquisition of plasmid transposon genes for drug resistance.
            • Genus: Peptostreptococcus
              • Features
                • Peptostreptococci are obligate anaerobic streptococci
                • They are part of the normal flora of the oral, intestinal, and genitourinary tracts
            • Genus: Bacillus
              • Features
                • Gram-positive, spore-forming rods that may form chains. Although spores survive for decades in dry environments, they quickly germinate in rich moist conditions (e.g., a macrophage) and transform into metabolically active vegetative cells.
                • They are aerobes (or facultative anaerobes) and grow well in ambient air.
                • They cause anthrax (Bacillus anthracis) and food poisoning (Bacillus cereus).
            • Genus: Clostridium. Common name: Clostridia
              • Features
                • Anaerobic, Gram-positive, spore-forming rods that can form chains.
                • Cause botulism (Cl. botulinum) characterized by a flaccid paralysis; gas gangrene and food poisoning (Cl. perfringens); and tetanus (Cl. tetani), characterized by rigid spasms.
            • Genus: Listeria. Listeria monocytogenes is the only human pathogen
              • Features
                • Short Gram-positive, non-spore-forming rod showing weak beta-hemolysis on blood agar.
                • Motile in broth by tumbling. (They lack forward movement.)
                • Facultative intracellular pathogens; they move from cell to cell by actin polymerization, which may propel the bacterium directly into an adjoining cell without exposure to extracellular milieu.
                • Grows in the cold and, unlike most non-spore-forming pathogens, survives in the environment.
                • Listeria monocytogenes is found in animal feces, rotting vegetation, and occasionally in soft cheeses, deli meats, and cabbage.
                • Causes mild gastroenteritis, as well as septicemia in pregnant women, leading to potential fetal septicemia or meningitis, and may cause meningitis in immunocompromised patients.
            • Genus: Erysipelothrix
              • Features
                • Aerobic Gram-positive, non-spore-forming rods.
                • Found in animals and rotting organic material; entry is through traumatic implantation.
                • Cause cutaneous erysipeloid primarily in fishmongers, butchers, and veterinarians.
            • Genus: Corynebacterium
              • Features
                • Club-shaped, Gram-positive non-spore-forming bacteria.
                • Found in Chinese character-like arrangement of cells.
                • Aerobic and nonmotile.
                • Part of normal flora; the non-toxin-producing corynebacteria found in the normal microbiota are called diphtheroids.
                • Tox+ C. diphtheriae causes diphtheria, and Corynebacterium jeikeium causes infections via catheters and foreign bodies in immunocompromised hosts.
            • Genus: Actinomyces
              • Features
                • Anaerobic, Gram-positive rods with some branching; nonmotile.
                • Found in crevices between teeth and gums and female genital tract.
                • Not acid-fast.
                  • These bacteria are related to mycobacteria.
                  • They have a similar cell wall but lack the extremely long chain fatty acids found in the mycobacterial cell wall. They have shorter fatty acid chains.
                • Cause cervicofacial or pelvic infections following trauma that has resulted in necrotic tissue; colonies formed in tissue are sometimes described as “sulfur” granules.
            • Genus: Nocardia.
              • Features
                • Gram-positive filamentous bacteria breaking up into rods.
                • Aerobic soil organism.
                • Often described as weakly or partially acid-fast as the slide will show some areas where the cells retain some of the carbol-fuchsin (hot pink red color).
                • Related to mycobacteria—have a cell wall with shorter chain mycolic acids; are somewhat resistant to drying, so they are transmitted in dust.
                • Cause tuberculosis-like (but not contagious) bronchopulmonary disease in immunocompromised patients.

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